Entomofauna of the reserve spruce forests is various. There are about 700 species of spruce vermin, and more than 600 species of them are insects. They inhabit weak, withering trees, and wind-fallen trees at the edges of the old wood-cutting areas and at the natural spruce windfall places. Most of the spruce vermin species are bark beetles (Scolytidae) and longhorn beetles (Cerambycidae).
Peacock butterfly (Inachis io)
Leucorrhinia intermedia
Grass eggar (Lasiocampa trifolii)
There are various butterflies in the reserve (more than 250 species). The most usual species are mourning cloak butterfly (Nymphalis antiopa), peacock butterfly (Inachis io), different species of Agrinnis, Colias, Lycaenidae. Looper Moth (Geometridae), Noctuidae, Phyllonorycter, Acronicta, leafrollers (Tortricidae) are also common everywhere. The fauna of predatory insects is also rich: sap beetles (Nitidulidae), hister beetles (Histeridae), flat bark beetles (Cucujidae). There are many parasite insects (Tachinid flies (Tachinidae), Ichneumonidae), which partly suppress the increasing in the numbers of longhorn beetles (Cerambycidae), bark beetles (Scolytidae) and other vermin. In the reserve were found 16 species of ants, 113 species of bugs, 23 species of horse-flies, 23 species of bumblebees and other insects.
Silver washed fritillary (Argynnis paphia)
Polygonia c – album .
Chalk-hill blue (Lysandra coridon)
In the reserve forests and on the adjacent protected territory (including areas thransformed by hunman activities) were found about 200 species of soil mesofauna: either permanent soil inhabitants (Myriapoda, worms, some beetles) or connected with soil only on larva stage (Diptera, beetles).
Oil beetle (Meloe proscarabaeus)
Longhorn beetle (Cerambycidae)
Common dor beetle (Geotrupes stercorarius)
Quite a few species, typical for the mixed and broad-leaved European forests, live only in nemoral spruce forests, and in the soils of the other types of spruce forests prevail widely spread species of taiga zone, which are ecologically connected with podzol and podzol-bog soils. Soil mesofauna of the spruce forests include several eurybiontic and synanthropic species. In the mesofauna prevail species with wide natural habitats - Holarctic, Palearctic and Eurosiberian species. There are less European species, except Diplopoda, who are mostly Eastern European forest species. There are few species from the Western Europe.
In the soils of the spruce forests prevail Diptera larvae, and beetle larvae and imago. These are the most various groups, and they dominate by numbers in the all types of spruce forests. The total amount of mesofauna in spruce forest soils changes depending on year and season. In the years, moderate by humidity and temperature, its numbers increase from relatively dry soils of nemoral spruce forests to wet soils of sedge-sphagnous spruce forests, but total mass and diversity decrease. For example, the number of species of Diplopoda, mollusca and earth-worms distinctly decrease even in spruce green-mossy fir forests.
By the type of foraging among invertebrates prevail predators, microphytophagans (which eat microscopical fungae and algae) and detritophagans. Invertebrates, which actively consume falled leaves (phytosaprophagans) prefer nemoral spruce forests, where they are more numerous and diverse. For phytosaprophagans it is difficult to access to the main mass of plant remains in boreal spruce forests, so there prevail microphytophagans and predators.
On the reserve territory and adjacent areas were found 162 spider species from 17 families and 70 genera. The most diverse are the families: Linyphiidae — 33 species, Areneidae — 20 species.
Raft spider (Dolomedes fimbriatus)
Wolfspider (Lycosidae)
Sheetweb weaver (Linyphiidae)
Spiders are more abundant in nemoral spruce forests — 68 species. There are 10-40 spiders per 1 square meter. In green-mosses spruce forests were found 44 spider species, in dwarf shrubs-sphagnum spruce forests - 42 species. Fauna of the sphagnous pine forests and peatbogs is poor - 13 species. Wood-cutting areas for the first five years are occupied by inhabitants of non-forest habitats, and during re-growing the composition of forest species gradually restore.
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